ISRA FACTSHEETS
ISRA FACTSHEETS
EUROPEAN ATLANTIC
Bermeo
Summary
Bermeo is located offshore of Vizcaya and Guipúzcoa provinces, Basque Country, Spain. This area encompasses part of the continental shelf and the slope towards the Capbreton canyon system. The habitat is characterised by rocky, sandy-muddy, and sandy substrates with pockmarks. The area is influenced by a marked seasonal stratification and mixing, with moderate wind-driven upwelling, eddies, and freshwater input from local rivers, contributing to nutrient enrichment and enhanced primary productivity. The area overlaps with the Cantabrian Sea (Southern Bay of Biscay) Ecologically or Biologically Significant Marine Area. Within this area there are: reproductive areas (Blue Shark Prionace glauca).
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Bermeo
DESCRIPTION OF HABITAT
Bermeo is located offshore of Vizcaya and Guipúzcoa provinces, Basque Country, Spain. This area encompasses part of the continental shelf and part of the slope towards the Capbreton canyon system. Bermeo is characterised by shallower areas on the continental shelf which are predominantly rocky and occasionally covered by sandy-muddy and sandy substrates, to the slope in which muddy-sandy or muddy sedimentation predominates with the presence of contourite channels with pockmarks (cone-shaped, circular, or elliptical depressions) (Jané et al. 2010; Galparsoro et al. 2020).
The area is influenced by a marked seasonal stratification and mixing, typical of temperate seas. During the boreal winter, the water column is fully mixed, whereas in spring and summer, surface warming leads to strong thermal stratification (Mason et al. 2006). The region also experiences moderate wind-driven upwelling and freshwater input from local rivers, contributing to nutrient enrichment and enhanced primary productivity (ICES 2022). In addition, the area is influenced by dynamic mesoscale features, including shelf-break fronts and eddies (Fernández et al. 2004; Borja et al. 2019). Submarine canyons along the continental slope off the Basque coast carry organic material downward, linking surface and deepwater ecosystems, and boosting local productivity and habitat diversity (van den Beld et al. 2017). Within the area, the anchovy Engraulis encrasicolus spawns seasonally from March to August, a process triggered by the warming of surface waters and the onset of water column stratification (Motos et al. 1996).
The area overlaps with the Cantabrian Sea (Southern Bay of Biscay) Ecologically or Biologically Significant Marine Area (EBSA; CBD 2025).
This Important Shark and Ray Area is pelagic and is delineated from inshore and surface waters (0 m) to 1,800 m based on the bathymetry of the area and the depth range of the Qualifying Species.
CRITERION C
SUB-CRITERION C1 – REPRODUCTIVE AREAS
Bermeo is an important reproductive area for one shark species.
Between 2003–2018, information was gathered from fisheries data from longline and purse seine observer programs in the Bay of Biscay (Druon et al. 2022). High capture rates of small-sized Blue Sharks (30–125 cm fork length [FL]) during July–December were reported from this area and surrounding waters. Size-at-birth for the species in the region is 39–41 cm FL (Skomal & Natanson 2003), with an estimated size of 95–100 cm FL for young-of-year (YOY) (Carlson et al. 2023), indicating that some of these individuals were neonates and YOY. Additionally, between 2020–2024, recreational snorkelling with Blue Sharks was conducted annually in the area and surrounding waters from June–October, averaging 15 trips per month. Chumming was used to attract the sharks to the surface. Between 2020–2024, Blue Sharks were observed during 194 shark snorkelling trips. From 2021–2024, a total of 17 individuals exhibited fresh mating scars, nine of which were confirmed to be females (C Barría et al. unpubl. data 2025). These individuals were sighted between July–October (C Barría et al. unpubl. data 2025). In 2023, one mating attempt was recorded in the area (L Vossgaetter pers. comm. 2025). Although Blue Sharks are widely distributed throughout the region, the regular and predictable presence of neonates, YOY, and mating behaviour highlights the importance of Bermeo as a reproductive area.
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